Christophe Baltzinger

Researcher on plants-ungulates interactions

Skill area


2006-2010 Boulanger Vincent. Herbivory pressure and vegetation communities’ dynamics: short and mid term influence of deer populations on plant diversity in forest ecosystems.  PhD co-supervised by Jean-Luc Dupouey, INRA, Champenoux and Sonia Saïd, ONCFS, Birieux. 

2009-2012 Chollet Simon. Highlighting and managing the deer effect: practical and theoretical lessons provided by the introduction of black-tailed deer to Haida Gwaii. PhD co-supervised by Jean-Louis Martin, Cefe-CNRS, Montpellier

2011-2014 Picard Mélanie. Consequences of endozoochorous transport on the composition of plant communities: a functional approach based on three ungulates. PhD co-supervised by Frédéric Gosselin, Irstea, Nogent-sur-Vernisson

2015-2019 Karimi Soroor. Endozoochorous Seed Dispersal by Mammalian Large Herbivores and Omnivores in Golestan National Park, Iran. PhD co-supervised by Dr. Mahmoud-Reza Hemami, Dr. Mostafa Tarkesh Esfahani and Dr. Hossein Akhani, Isfahan University, Iran

2016-ongoing Shukla Ushma. Epizoochorous seed dispersal by wild ungulates in changing landscapes – the red deer as a case study. PhD co-supervised by Björn Reineking, Irstea, Grenoble

2005-2007 Adrados Christophe. How to predict bedsite selection by Cervus elaphus using GPS locations and activity records? Post-doc co-supervised by Dominique Pépin, INRA, Castanet-Tolosan.

2010-2011 Pellerin Maryline. Long-distance plant dispersal by a guild of wild ungulates. Post-doc co-supervised by Sonia Saïd and Eric Baubet, ONCFS, Birieux

2012-2014 Albert Aurélie. How to predict the role of wild ungulates in long-distance plant dispersal in temperate ecosystems using a comparative approach based on functional traits? Post-doc co-supervised by Anders Mårell, Irstea, Nogent-sur-Vernisson.

And 15 Master of Science students from 2005 up to now

Ongoing and past projects

  • 2009-2015 DIPLO 1 & 2 Long-distance Plant Dispersal by wild Ungulates (Loiret, France)
  • 2012-2015 DYSPERSE Spatio-temporal fencing dynamics in Sologne and Ecosystem Services (Sologne, France)
  • 2014-2016 DUNG BEETLE Dung removal and secondary seed dispersal by dung beetles along a biogeographic and climatic gradient throughout the Western Palaearctic
  • 2015-2018 COSTAUD Contribution of wild ungulates to ecosystem functioning and cultural services in Chambord (Loir-et-Cher, France)
  • 2005-2010 ARC EN BARROIS Short and mid-term effects of deer populations on forest understory plant communities (Haute-Marne, France)
  • 2008-2014 GNB Management, Naturalness and Biodiversity (France)
  • 1998-2003 BOUGES Bedsite selection by sympatric red and roe deer (Lozère, France)
  • 1997-2015 HAIDA GWAII Restoring forest ecosystem functioning after drastic reduction of black-tailed deer abundance (British Columbia, Canada)


Group of interest: EURODEER

Associate scientist: Research Group on Introduced Species (RGIS), Forest Ungulate Research Network (FURN), Silviculture and Ungulates WP (IUFRO), Frugivores and Seed Dispersal (FSD2020) and ALTER-Net

ONCFS Hunting and Wildlife National Agency Unit Wild ungulates
INRA Research Units Wildlife Ecology and Behaviour, Castanet-Tolosan & Forest Ecology and Ecophysiology, Nancy, France
ENVA (Alfort National Veterinary School), Champignelles, France
CEFE-CNRS (Dr. Jean-Louis Martin), Montpellier, France

Contributor to peer-reviewed journals (Publons)




Google Scholar profile - Orcid - Rg - ResearcherID  and from 2015 ...


  • Karimi, S., Hemami, M. R., Esfahani, M. T. & Baltzinger, C. (in revision) Potential and realised endozoochorous dispersal by herbivores and omnivores in a Caspian forest steppe area. Perspectives in Plant Ecology, Evolution and Systematics


  • Baltzinger, C., Karimi, S. & Shukla, U. 2019. Plants on the move: hitch-hiking with ungulates distributes diaspores across landscapes. Frontiers in Ecology and Evolution link

  • Cocquelet, A., Mårell, A., Bonthoux, S., Baltzinger, C. & Archaux, F. 2019. Direct and indirect effects of ungulates on forest birds' nesting failure? An experimental test with artificial nests. Forest Ecology and Management 437(1): 148-155 link

  • Milotić, T., Baltzinger, C., Eichberg, C., Eycott, A., Heurich, M., Müller, J., . . & Hoffmann, M. 2019. Functionally complete communities results in better ecosystem functioning: Dung removal and secondary seed dispersal by dung beetles in the Western Palaearctic. Journal of Biogeography link

  • Milotić, T., Baltzinger, C., Eichberg, C., Eycott, A., Heurich, M., Müller, J., . . & Hoffmann, M. 2018. Dung beetle assemblages, dung removal and secondary seed dispersal: data from a large-scale, multi-site experiment in the Western Palearctic. Frontiers of Biogeography 10.1, e37829 link

  • Karimi, S., Hemami, M. R., Esfahani, M. T. & Baltzinger, C. 2018. The Role of Brown Bear (Ursus arctos) in the Plant Seed Dispersal of Golestan National Park. Iranian Journal of Applied Ecology link

  • Karimi, S., Hemami, M. R., Esfahani, M. T., Akhani, H. & Baltzinger, C. 2018. Complementary endozoochorous seed dispersal by large mammals in the Golestan National Park, Iran. Seed Science Research link

  • Liehrmann, O., Jégoux, F., Guilbert, M.A.,Isselin-Nondedeu, F., Saïd, S., Locatelli, Y. & Baltzinger, C. 2018. Epizoochorous dispersal by ungulates depends on fur, grooming and social interactions. Ecology and Evolution 8(3): 1582-1594 link

  • Boulanger, V., Dupouey, J.L., Archaux, F., Badeau, V., Baltzinger, C., Chevalier, R., Corcket, E., Dumas, Y., Forgeard, F., Mårell, A., Montpied, P., Paillet, Y., Picard, J.-F., Saïd, S. & Ulrich E. 2018. The role of ungulates in nowadays temperate forests. A response to Fløjgaard et al. (DOI:10.1111/gcb.14029). Global Change Biology link

  • Boulanger, V., Dupouey, J.L., Archaux, F., Badeau, V., Baltzinger, C., Chevalier, R., Corcket, E., Dumas, Y., Forgeard, F., Mårell, A., Montpied, P., Paillet, Y., Picard, J.-F., Saïd, S. & Ulrich E. 2018. Ungulates increase forest plant species richness to the benefit of non-forest specialists. Global Change Biology 24(2): e485–e495 link

  • Lalleroni, A., Quenette, P.-Y., Daufresne, T., Pellerin, M., & Baltzinger, C. 2017. Exploring the potential of brown bear (Ursus arctos arctos) as a long-distance seed disperser: a pilot study in South-Western Europe. Mammalia 81(1): 1-9 link

  • Picard, M., Chevalier, R., Barrier, R., Boscardin, Y. & Baltzinger, C. 2016. Functional traits of seeds dispersed through endozoochory by native forest ungulates. Journal of Vegetation Science 27(5): 987–998 link

  • Pellerin, M., Picard, M., Saïd, S., Baubet, E. & Baltzinger, C. 2016. Complementary endozoochorous long-distance seed dispersal by three native herbivorous ungulates. Basic and Applied Ecology 17(4): 321-332 link

  • Albert, A., Auffret, A., Cosyns, E., Cousins, S., D'hondt, B., Eichberg, C., Eycott, A., Heinken, T., Hoffmann, M., Jaroszewicz, B., Malo, J., Mårell, A., Mouissie, M., Pakeman, R., Picard, M., Plue, J., Poschlod, P., Provoost, S., Schulze, K.A. & Baltzinger, C. 2015. Seed dispersal by ungulates as an ecological filter : a trait-based meta-analysis. Oikos 124(9): 1109-1120 link

  • Picard, M., Papaïx, J., Gosselin, F., Picot, D., Bideau, E. & Baltzinger, C. 2015. Temporal dynamics of seed excretion by wild ungulates: implications for plant dispersal. Ecology and Evolution 5(13): 2621-2632 link

  • Albert, A., Mårell, A., Picard, M. & Baltzinger, C. 2015. Using basic plant traits to predict ungulate seed dispersal potential. Ecography 38(5): 440-449 link

  • Boulanger, V., Baltzinger, C., Saïd, S., Ballon, P., Picard, J.-F. & Dupouey, J.-L. 2015. Decreasing deer browsing pressure influenced understory vegetation dynamics over 30 years. Annals of Forest Science 72(3): 367-378 link


Irstea, Forest Ecosystems Research Unit

Domaine des Barres

45290 Nogent-sur-Vernisson, France
+ 33 (0)2 38 95 03 59
+33 (0)2 38 95 66 75